冰灾干扰对粤北常绿阔叶林群落特征、空间结构和群落稳定性的中长期影响OA
Medium to Long Term Effects of Ice Storm Disturbance on Community Characteristics,Spatial Structure,and Community Stability of Evergreen Broad-Leaved Forests in Northern Guangdong,China
[目的]对比分析粤北常绿阔叶林受冰灾干扰 13年后与未受冰灾干扰森林在群落特征、空间结构和群落稳定性方面的差异,为粤北森林的生物多样性保护和可持续发展提供决策依据.[方法]在粤北南雄市北部帽子峰、南部青嶂山分别建立 1块 1 hm2 遭受冰灾干扰的常绿阔叶林大样地和 1块未受冰灾干扰的对照大样地,对大样地内胸径>1 cm的所有树木进行每木检尺,记录树种名、胸径、树高和相对坐标等信息.对比分析遭受和未受冰灾干扰的常绿阔叶林在树种组成、生物多样性指数、林分径阶和树高分布等方面的差异,采用相邻木构成的空间结构单元方法分析常绿阔叶林的角尺度、大小比数和平均混交度 3个林分空间结构指标,应用Godron 方法分析常绿阔叶林的群落稳定性.[结果]遭受和未受冰灾干扰的大样地均为以壳斗科为主要优势种的典型亚热带常绿阔叶林,遭受冰灾干扰大样地内重要值排名前三的优势种为壳斗科的栲、米槠和罗浮锥,未受冰灾干扰大样地内重要值排名前三的优势种为罗浮锥、木荷和青冈.遭受冰灾干扰大样地内胸径为 1~4 cm的小径阶树木数量(2 668株)远高于未受冰灾干扰大样地(681株),遭受和未受冰灾干扰大样地内胸径为 4~30 cm的中等径阶树木数量分别为 1 441和 1 345株,胸径>30 cm的大径阶树木数量分别为 79和 82株,二者差异不大.遭受和未受冰灾干扰大样地内树木的平均角尺度分别为 0.56和 0.62,说明随机分布是两样地中大多数树木的主要分布方式,但遭受冰灾干扰样地部分种群有呈聚集分布演替趋势.两样地树木的大小比数均值分别为 0.51和 0.49,各等级占比相差很小,表明树木优劣分化程度不大.两样地树木的平均混交度分别为 0.79和 0.62,表明树种间的空间隔离程度很高,且遭受冰灾干扰样地的林分混交度优于未受冰灾干扰样地.遭受冰灾干扰样地的群落稳定性与理想的稳定森林群落存在一定差距,而未受冰灾干扰样地森林群落与理想的稳定森林群落非常接近.[结论]冰灾干扰对粤北常绿阔叶林的群落特征、林分空间结构和群落稳定性均有较大影响,即使在冰灾干扰后已恢复 13年,但遭受冰灾干扰的常绿阔叶林内小树比例仍然较高,林分密度仍然较大,群落中部分种群有呈聚集分布趋势,群落稳定性仍然较低.冰灾干扰对粤北常绿阔叶林具有长久影响,受损森林恢复将是一个长期的动态过程.
[Objective]This study aims to investigate the medium to long term effects of ice storm disturbance on natural evergreen broad-leaved forests,and to clarify the differences in community characteristics,stand spatial structure,and community stability between ice-damaged forests after 13 years of ice storm disturbance and undisturbed in northern Guangdong Province,China.[Method]One 1 hm2 permanent plot affected by the 2008 ice storm was established in Maozifeng(northern Nanxiong),and a comparable 1 hm2 undisturbed control plot was established in Qingzhangshan(southern Nanxiong).All trees with a diameter at breast height(DBH)≥1 cm were recorded and measured,including species identity,DBH,tree height,and spatial coordinates.Comparative analysis was conducted on the differences in species composition,biodiversity indices,DBH,and height class distributions between the two plots.The neighborhood-based spatial structural unit method was used to analyze the stand spatial structure,including the uniform angle index,neighborhood comparison,and mingling.The Godron method was used to analyze community stability.[Result]The two plots affected and unaffected by ice disasters represented typical subtropical evergreen broad-leaved forests dominated by Fagaceae species.In the ice-damaged plot,the three dominant species(by importance value)were Castanopsis fargesii,C.carlesii,and C.faberi,whereas in the undisturbed plot they were C.faberi,Schima superba,and Quercus glauca.The number of small-diameter trees(DBH 1-4 cm)in the ice-damaged plot(2 668 individuals)was much higher than that in the undisturbed plot(681 individuals),while the numbers of medium-diameter trees(DBH 4-30 cm)and large-diameter trees(DBH>30 cm)were similar between the two plots.The mean uniform angle index of trees in the large sample plots affected and unaffected by ice disasters was 0.56 and 0.62,respectively,indicating that random distribution was the predominant spatial pattern in both plots,although some populations in the ice-damaged plot showed a tendency toward aggregated distribution.Mean neighborhood comparison values(0.51 and 0.49,respectively)suggested weak size differentiation and relatively stable stand structures.The mean mingling was higher in the ice-damaged plot(0.79)than in the undisturbed plot(0.62),indicating stronger interspecific mixing following disturbance.Community stability in the ice-damaged plot remained lower than that of an ideal stable forest,whereas the undisturbed plot closely approximated the stability of an ideal forest community.[Conclusion]Ice storm disturbance has pronounced and lasting effects on community characteristics,stand spatial structure,and community stability of evergreen broad-leaved forests in northern Guangdong.Even 13 years after the ice storm,the disturbed forest still exhibits a high proportion of small trees,elevated stand density,partial aggregation of tree populations,and reduced community stability.These results indicate that ice storm disturbance exerts long-term influences on subtropical evergreen broad-leaved forests,and that post-disturbance forest recovery is a prolonged and dynamic process.
邱治军;胡辉;郭绪兵;赵厚本;李兆佳
中国林业科学研究院热带林业研究所 广州 510520||南岭北江源森林生态系统定位观测研究站 广州 510520广东南雄小流坑-青嶂山省级自然保护区管理处 南雄 512400广东南雄小流坑-青嶂山省级自然保护区管理处 南雄 512400中国林业科学研究院热带林业研究所 广州 510520||南岭北江源森林生态系统定位观测研究站 广州 510520中国林业科学研究院热带林业研究所 广州 510520||南岭北江源森林生态系统定位观测研究站 广州 510520
农业科技
群落稳定性常绿阔叶林角尺度大小比数混交度
community stabilityevergreen broad-leaved forestuniform angle indexneighborhood comparisonmingling
《林业科学》 2026 (4)
45-54,10
中国林业科学研究院基本科研业务费专项(CAFYBB2021SY002)国家林业和草原局林业科技创新平台运行补助项目(2020132015).
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